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88  Lamiaceae (Draft: R. Elven)

S Labiatae



THIS DRAFT HAS NOT BEEN DISCUSSED

Comments:

(1) This large family is virtually absent from the Arctic except for a slight penetration along the southern boundary, and for one genus: Thymus. This lack is also relevant for most antropochores and I have been strict in acceptance of taxa as stable. A few more are temporarily mentioned in comments. (Elven)

88.1  Ajuga L. (1753), Sp. Pl. 561.
88.1.1  Ajuga pyramidalis L. (1753), Sp. Pl. 561.

S

2n= 32 (4x).



2nD Löve & Löve (1975) list seven counts, one Icelandic.

G NOR*


Comments:

(1) Reaches the Arctic along the Varangerfjord in NE Norway, as native or more probably as a stable archaeophyte. (Elven)



88.2  Scutellaria L. (1753), Sp. Pl. 598.
88.2.1  Scutellaria galericulata L. (1753), Sp. Pl. 599.

S

2n= 28-32.



2nD Gadella & Kliphuis (1968 W Eur, 2n=31, 32); Lövkvist & Hultgård (1999 Sweden, 2n=28, 30).

G RUS? CAN

Comments:

(1) If var. pubescens Benth. *** (Hultén 1968, Porsild & Cody 1980) is accepted it could be treated as an American subspecies due to its wide range. (Elven)



88.3  Glechoma L. (1753), Sp. Pl. 578.
88.3.1  Glechoma hederacea L. (1753), Sp. Pl. 578.

S

2n= (1) 18 (2x). (2) 36 (4x).



2nD (1) Skalinska et al. (1959 Poland); Sorsa (1963 Finl.). (2) Skalinska et al. (1959 Poland); Lövkvist & Hultgård (1999 Sweden, 31 counts from 15 sites).

G RUS SIB

Comments:

(1) As to Siberia: Given by Tolmachev & Yurtsev (1980) from lower Ob and a site mapped here by Hultén & Fries (1986) is inside the Arctic. (Elven)



88.4  Dracocephalum L. (1753), Sp. Pl. 594.
88.4.1  Dracocephalum palmatum Steph. ex Willd. (1800), Sp. Pl. 3: 151.

S

2n= 12 (2x).



2nD Zhukova (1967a).

G SIB RFE

Comments:
88.4.2  Dracocephalum parviflorum Nutt. (1818), Gen. N. Amer. Pl. 2: 35.

S

2n=



2nD

G CAN*?


Comments:

(1) Mapped by Porsild & Cody (1980) from one point on the SW shore of Hudson Bay ('Baffin-Hudson'), by Scoggan (1979) given as Churchill (as Molvadica parviflora (Nutt.) Britt.). Canadians must decide whether it is native, and if not whether it is stable. (Elven)



WARNING! Will probably be excluded either as non-arctic or as non-stable.

88.5  Prunella L. (1753), Sp. Pl. 600.
88.5.1  Prunella vulgaris L. (1753), Sp. Pl. 600.

S

Comments:



(1) Hultén (1968) accepts the American plants as a separate subspecies. This approach seems well justified and is followed here. (Elven)
88.5.1.1  Prunella vulgaris L. subsp. vulgaris

S

2n= 28 (4x).



2nD Löve & Löve (1975) list numerous counts, two Icelandic; Lövkvist & Hultgård (1999 Icel & Sweden).

G ICE? NOR*? RUS

Commnts:

(1) A border case in Iceland acc. to Kristinsson's map and list. Occurrence in arctic mainland Norway to be checked; here obviously introduced but probably a stable archaeophyte. (Elven)


88.5.1.2  Prunella vulgaris L. subsp. lanceolata (Barton) Hultén (1949), Lunds Univ. Årsskr., n. f., avd. 2, 45, 1: 1364.

B P. pensylvanica Willd. var. (beta) lanceolata Barton (1818), Comp. Fl. Philad. 2: 37.

2n= 28 (4x)

2nD Löve & Löve (1975) list some non-arctic American counts for the collective species, i.e., this subspecies.

G CAN?

Comments:



(1) A border case in the Hudson Bay area ('Baffin-Hudson'). Scoggan (1979) reports it north to Fawn River at 5440'N. (Elven)

WARNING! Might be excluded as non-arctic.

88.6  Galeopsis L. (1753), Sp. Pl. 579.

Comments:

(1) Two other species occur in the boundary areas, probably only as casuals: G. speciosa Miller (1768), Gard. Dict., ed. 8 , no. 3, 2n=16, in Polar Ural; and G. ladanum L. (1753), Sp. Pl. 579, 2n=16, in Polar Ural. (Elven)
88.6.1  Galeopsis bifida Boenn. (1824), Prodr. Fl. Monast. Westphal. 178.

S

2n= 32 (4x).



2nD Müntzing (1929, C & SE Eur, Sweden).

G NOR RUS*? CAN*?

Comments:

(1) Fairly regular in maritime driftwall vegetation into the arctic parts in N Norway and must therefore be considered as native or archeophytic. Much more dubious as stable in other areas. (Elven)


88.6.2  Galeopsis tetrahit L. (1753), Sp. Pl. 579.

S

2n= 32 (4x).



2nD Müntzing (1929 W Eur, Sweden); Lövkvist & Hultgård (1999 Sweden).

G NOR* RUS*?

Comments:

(1) Much less bound to seashores than the previous species and more casual in the northern parts but still stable in arctic N Norway. (Elven)



88.7  Lamium L. (1753), Sp. Pl. 579.

Comments:

(1) Several other species occur, at least as casuals, in the southern border areas: L. amplexicaule L. (1753), Sp. Pl. 579, SW Greenland; L. confertum Fr. (1846), Summa Veg. Scand. 15, SW Greenland; and L. hybridum Vill. (1786), Hist. Pl. Dauphiné 1: 251, N Norway. (Elven)
88.7.1  Lamium album L. (1753), Sp. Pl. 579.

S

2n= 18 (2x).



2nD Pólya (1949 C Eur).

G ICE* RUS SIB

Comments:
88.7.2  Lamium purpureum L. (1753), Sp. Pl. 579.

S

2n= 18 (2x).



2nD Sorsa (1963 Finl.).

G ICE* RUS**? GRL**

Comments:

(1) A casual in several areas but reported by Kristinsson as stable in N Iceland. Therfore included. (Elven)



88.8  Stachys L. (1753), Sp. Pl. 580.
88.8.1  Stachys palustris L. (1753), Sp. Pl. 580.

S

Comments:



(1) The American plants are recognised as a race and should probably be treated as a subspecies according to our criteria. The Eurasiatic subsp. palustris does not seem to reach the Arctic. (Elven)
88.8.1.1  Stachys palustris L. subsp. pilosa (Nutt.) Epling (1934), Feddes Repert. Beih. 80: 63.

B Stachys taxon (*) pilosa Nutt. (1834), J. Acad. Philad. 7: 48.

S

2n=


2nD

G CAN


Comments:

88.9  Lycopus L. (1753), Sp. Pl. 21.
88.9.1  Lycopus uniflorus Michx. (1803), Fl. Bor.-Amer. 1: 14.

S

2n=



2nD

G CAN


Comments:

(1) Seems to reach the Arctic at the SE side of Hudson Bay. (Elven)



88.10  Mentha L. (1753), Sp. Pl. 576.

Comments:

(1) Both the entities listed below belong to a M. arvensis aggregate. (Elven)
88.10.1  Mentha arvensis L. (1753), Sp. Pl. 577.

S

2n= (1) 24. (2) 72. (3) 90.



2nD (1) Ouweneel (1968 W Eur); (2) Olsson (1967 Sweden), Lövkvist & Hultgård (1999 Sweden); (3) Ouweneel (W Eur).

G RUS RFE ALA CAN

Comments:

(1) Hultén (1968) and Porsild & Cody (1980) includes a var. villosa (Benth.) Stewart (1944), Rhodora 46: 331, for the American parts. However, the species is very polymorphic and many geographical varieties (e.g. var. lapponica ***) have been described. The occurrences mapped in European Russia probably belong to var. lapponica, those from N/C North America to var. villosa, and those from Alaska and Russian Far East to some other. I can only support Hultén (1949 p. 1368): "In Alaska, as in most other places, this species occurs in several different forms. It seems to me to be of no value to try to report them under separate names ...". (Elven)


88.10.2  Mentha sachalinensis (Briq.) Kudo (1921), J. Coll. Sci. Univ. Tokyo 43, 8: 47.

B M. arvensis L. var. sachalinensis Briq. ***

S

2n= 100.


2nD Sokolovskaya (1960a).

G RFE?


Comments:

(1) Relations to M. arvensis must be clear before acceptance. The single chromosome count might indicate a higher ploidy level but the levels reported for M. arvensis (and other species of the genus) vary widely. (Elven)



WARNING! Might be excluded as non-arctic or merged with M. arvensis s. lat.

88.11  Thymus L. (1753), Sp. Pl. 590.

Comments:

(1) Menitsky in Tolmachev & Yurtsev (1980, Fl. Arct. URSS 8) also includes a T. ochotensis Klok. (1954), Bot. Mater. Gerb. Bot. Inst. AN SSSR 16: 300, from lower Kolyma (SIB/RFE). Arctic? (Elven)

(2) Species concepts vary much in this genus. In N Europe there is a tradition that follows J. Jalas and accepts a few wide species (esp. T. praecox and T. serpyllum) with several geographical races. In C and S Europe and Russia more narrowly circumscribed species are often applied. This influences the arctic treatmentas Menitsky follows this second tradition.

The only distinctly arctic taxon of the entire family ('arcticus') is treated as a subspecies of the W/C European T. praecox or alternatively of the related T. polytrichus. The N Fennoscandian entity ('tanaensis') is treated as a subspecies of the C/E European and Siberian T. serpyllum. The Russian entities are uniformly treated as species even if they probably are less clearly separated from each other and from T. serpyllum than is 'tanaensis'. This imbalance in treatments must be solved. As all the disputed taxa are restricted to Russia   Siberia, we need good Russian input here. They must also consider whether these Russian taxa are more or less distinct than the N Atlantic and N European taxa currently treated as subspecies. (Elven)
88.11.1  Thymus praecox Opiz (1824), Naturalientausch 6: 40.

S

Comments:


88.11.1.1  Thymus praecox Opiz subsp. arcticus (E. Durand) Jalas (1970), Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 43: 190.

B T. serpyllum L. var. arcticus E. Durand in Kane (1856), Arct. Explor. 1853-55, App. 18: 459.

S T. arcticus (E. Durand) Ronn. (1924), Feddes Repert. 20: 331, 237; T. drucei Ronn. (1924), Feddes Repert. 20: 328; T. polytrichus A. Kern. ex Borbás [(1890), Természettud. Közl. 24: 105] subsp. britannicus (Ronn.) Kerguélen (1887), Lejeunia, n. s., 120: 175; T. praecox Opiz subsp. britannicus (Ronn.) Holub (1973), Preslia 45: 359.

2n= c.50-54.

2nD Pigott (1954 W Eur, 2n=c.50, 51, 54); Jørgensen et al. (1958 Grl, 2n=54).

G ICE GRL

Comments:

(1) The ambiguity as to which species the arctic plants belong, T. praecox (1824) or T. polytrichus (1890), might justify treatment as a separate species. I am uncertain which of Ronnigers names   drucei or arcticus - would have priority. (Elven)


The Thymus serpyllum L. aggregate (T. extremus, T. oxyodontus, T. pauciflorus, T. reverdattoanus, T. serpyllum)

Comments:

(1) The T. reverdattoanus aggregate of Menitsky in Tolmachev & Yurtsev (1980, Fl. Arct. URSS 8) seems to be closely related to T. serpyllum, i.e., parts of an extended T. serpyllum aggregate. Thymus serpyllum subsp. tanaensis then seems to be the westernmost of a series of more or less defined races (or species). The treatment of 'tanaensis' as a subspecies of T. serpyllum and the others as separate species is probably a difference in taxonomic traditions, not in biology. This must be 'corrected' in the next version of the checklist draft. (Elven)
88.11.2  Thymus serpyllum L. (1753), Sp. Pl. 590.

S

Comments:


88.11.2.1  Thymus serpyllum L. subsp. tanaensis (Hyl.) Jalas (1947), Acta Bot. Fenn. 39: 20.

B T. serpyllum L. var. tanaensis Hyl. (1945), Uppsala Univ. Årsskr. 1945, 7: 276.

S

2n= 24.


2nD Jalas (1948 Finl.).

G NOR


Comments:

(1) Reaches the Arctic in N Norway in Vadsø and approaches in Tana. This entity seems to be as well or better differentiated from T. serpyllum s. str. than the Russian entities described as species. I am not sure whether it ever has been described at level of species, but I believe not. (Elven)


88.11.3  Thymus pauciflorus Klok. (1954), Bot. Mater. Gerb. Bot. Inst. AN SSSR 16: 311.

S

2n=



2nD

G RUS


Comments:

(1) Jalas (1972, Fl. Eur. 3) includes this species in T. talijevi Klok. & Schost. (1936), J. Inst. Bot. Acad. Sci. Ukr. 9: 195, and he writes that it is close to T. serpyllum subsp. tanaensis. (Elven)


88.11.4  Thymus reverdattoanus Serg. (1936), Sist. Zam. Gerb. Tomsk. Univ. 1-2: 5.

S

2n=



2nD

G SIB


Comments:
88.11.5  Thymus extremus Klok. (1954), Bot. Mater. Gerb. Bot. Inst. AN SSSR 16: 308.

S

2n=



2nD

G RUS? SIB

Comments:
88.11.6  Thymus oxyodontus Klok. (1954), Bot. Mater. Gerb. Bot. Inst. AN SSSR 16: 311.

S

2n=



2nD

G SIB RFE



Comments:


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