Nbaf complete bibliography Please note that the list of publications for 2016 will be completed early 2017

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  1. Azadi NA (2007) MSc Thesis (Lancaster University) Microarray analysis of the effect of industrial pollution on stickleback fish using linear mixed effect models.

  2. Bamford A (2007) PhD Thesis (University of Nottingham) Competition and co-existence among vultures: from individual behaviour to conservation ecology.

  3. Berg M (2007) PhD Thesis (University of Groningen) Sexual selection and reproductive strategies in songbirds : territoriality, mate and parental care.

  4. Bourke B (2007) PhD Thesis (University of Nottingham). Population genetics of the golden eagle (Aquila chrysaetos) in the British Isles and its phylogeography in Europe.

  5. *Coiffait L (2007) PhD Thesis (Newcastle University). Investigating the movements of migratory thrushes Turdus merula and T. iliacus using intrinsic markers and morphology.

  6. *Dawson DA (2007) PhD Thesis (University of Sheffield) Genomic analysis of passerine birds using conserved microsatellite loci.

  7. Dugdale H (2007) PhD Thesis (University of Oxford). The evolution of social behaviour: the effect of mating system and social structure in the European badger Meles meles.

  8. Geoghegan F (2007) PhD Thesis (The University of Birmingham – NBAF-E) Differential gene expression in the three-spined stickleback, Gasterosteus aculeatus, following exposure to mixtures of compounds.

  9. Li W (2007) PhD Thesis (University of Liverpool). An integrated suite of informatics tools and resources to support post-genomics investigations.

  10. Mainwaring MC (2007) PhD thesis (Lancaster University). Evolutionary causes and consequences of hatching asynchrony in nestling birds.

  11. Maunder R.J. (2007) PhD Thesis (CEH Lancaster, Lancaster Environment Centre - NBAF-L) The effects of endocrine disrupting chemicals on reproductive performance of sticklebacks.

  12. O´Connor E. (2007) PhD Thesis (University of Liverpool - NBAF-L) Population difference in response to hypoxia: Behaviour and physiology of three-spined stickleback, Gasterosteus aculeatus.


[NBAF-E] 8

  1. Cutter AD, Wasmuth JD, Blaxter ML (2006) The evolution of biased codon and amino acid usage in nematode genomes. Mol Biol Evol 23 2303-2315.

  2. Dalmay T (2006) Short RNAs in environmental adaptation. Proc R Soc Lond B 273 1579-1585.

  3. Dalmay T, Edwards DR (2006) MicroRNAs and the hallmarks of cancer. Oncogene 25 6170-6175.

  4. Hegarty MJ, Barker GL, Wilson ID, Abbott RJ, Edwards KJ, Hiscock SJ (2006) Transcriptome shock after interspecific hybridization in Senecio is ameliorated by genome duplication. Curr Biol 16 1652-1659.

  5. Ho T, Pallett D, Rusholme R, Dalmay T, Wang H (2006) A simplified method for cloning of short interfering RNAs from Brassica juncea infected with turnip mosaic potyvirus and turnip crinkle carmovirus. J Virol Methods 136 217-223.

  6. Jones M. & Blaxter M (2006) TaxMan: A taxonomic database manager. BMC Bioinformatics 7 536-539. [T]

  7. Rathjen T, Nicol C, McConkey G, Dalmay T (2006) Analysis of short RNAs in the malaria parasite and its red blood cell host. FEBS Lett 580 5185-5188.

  8. Zaldivar-Riveron A, Mori M, Quicke DL (2006) Systematics of the cyclostome subfamilies of braconid parasitic wasps (Hymenoptera: Ichneumonoidea): a simultaneous molecular and morphological Bayesian approach. Mol Phylogenet Evol 38 130-145.

[NBAF-L] 9

  1. Allen MJ, Wilson WH (2006) The coccolithovirus microarray: an array of uses. Briefings in Functional Genomics and Proteomics 5 273-9

  2. Chipman JK, George SG, Williams TD, Diab AM, Sabine V, Ortega F and Falciani F (2006) A high density flounder (Platichthys flesus) cDNA microarray as a tool in the identification of expression changes in gene sets predictive of exposure to pollutants. Toxicology Letters, 164 Supplement 1, S42 [not referred].

  3. Cossins AR, Fraser EJ Gracey AY (2006) Post-genomic approaches to understanding the mechanisms of environmentally-induced phenotypic plasticity. J. exp. Biol. 209 2328-2336.

  4. Fraser J, Mello LV, Fang Y, Brass A, Gracey AY, Cossins AR (2006) Hypoxia-inducible myoglobin expression in non-muscle tissues. Proc. Nat. Acad. Sci. USA 103 2977-2981.

  5. Lewis NA, Williams TD, Chipman JK (2006) Functional analysis of a metal response element in the regulatory region of flounder cytochrome P450 1A and implications for environmental monitoring of pollutants. Toxicol Sci 92 387-393.

  6. µMorrison N, Wood AJ, Hancock D, Shah S, Hakes L, Gray T, Tiwari B, Kille P, Cossins A, Hegarty M, Allen MJ, Wilson WH, Olive P, Last K, Kramer C, Bailhache T, Reeves J, Pallett D, Warne J, Nashar K, Parkinson H, Sansone SA, Rocca-Serra P, Stevens R, Snape J, Brass A, Field D.§ (2006) Annotation of environmental OMICS data: application to the transcriptomics domain. ‘OMICS 10 172-8.

  7. Sheader DL, Williams TD, Lyons BP, Chipman JK (2006) Oxidative stress response of European flounder (Platichthys flesus) to cadmium determined by a custom cDNA microarray. Mar Environ Res 62 33-44.

  8. Williams TD, Diab AM, George SG, Godfrey RE, Sabine V, Conesa A, Minchin SD, Watts PC, Chipman JK (2006) Development of the GENIPOL European flounder (Platichthys flesus) microarray and determination of temporal transcriptional responses to cadmium at low dose. Environ Sci Technol 40 6479-6488.

  9. Wilson R, Cossins AR, Spiller DG (2006) Encoded microcarriers for high-throughput multiplexed detection. Angewandte Chemie-International Edition 45 6104-6117.

[NBAF-S] 12

  1. Anderson SJ, Dawson DA, Freeland JR (2006) Isolation and characterisation of highly polymorphic microsatellite loci for the garden tiger moth Arctia caja (Lepidoptera: Arctiidae). Molecular Ecology Notes, 6, 104-106.

  2. Bearhop S, Phillips RA, McGill R, Cherel Y, Dawson DA, Croxall JP (2006) Stable isotopes indicate sex-specific and long-term individuals foraging specialisation in diving seabirds. Marine Ecology Progress Series, 311, 157-164.

  3. Bourke B, Dawson DA (2006) Fifteen microsatellite loci characterised in the golden eagle Aquila chrysaetos (Accipitridae: Aves). Molecular Ecology Notes, 6, 1047-1050.

  4. Chen SF, Rossiter SJ, Faulkes CJ, Jones G (2006) Population genetic structure and demographic history of the endemic Formosan lesser horseshoe bat (Rhinolophus monoceros) Molecular Ecology, 15, 1643–1656.

  5. *Dawson DA, Burke T, Hansson B, Pandhal J, Hale MC, Hinten GN, Slate J (2006) A predicted microsatellite map of the passerine genome based on chicken-passerine sequence similarity. Molecular Ecology, 15, 1299-1320.

  6. Frantz AC, Fack F, Muller CP, Roper TJ (2006) Faecal DNA typing as a tool for investigating territorial behaviour of badgers (Meles meles). European Journal of Wildlife Research, 52, 138-141.

  7. Frantz AC, Roper TJ (2006) Simulations to assess the performance of different rarefaction methods in estimating population size using small datasets. Conservation Genetics, 7, 315-318.

  8. Hamill RM, Noor MAF, Watson TE, Ritchie MG (2006) New microsatellite loci for the European bushcricket Ephippiger ephippiger (Orthoptera: Tettigoniidae). Molecular Ecology Notes, 6, 340-342.

  9. Johnson PCD, Webster LMI, Adam A, Buckland R, Dawson DA, Keller LF (2006) Abundant variation in microsatellites of the parasitic nematode, Trichostrongylus tenuis and linkage to a tandem repeat. Molecular and Biochemical Parasitology, 148, 210-218.

  10. Leitner S, Marshall RC, Leisler B, Catchpole CK (2006) Male song quality, egg size and offspring sex in captive canaries, Serinus canaria. Ethology, 112, 554-563.

  11. Rivers NM, Butlin RK, Altringham JD (2006) Autumn swarming behaviour of Natterer's bats in the UK: Population size, catchment area and dispersal. Biological Conservation, 127, 215-226.

  12. Rossiter SJ, Faulkes CG, Ransome RD, Faulkes CG, Dawson DA, Jones G (2006) Long-term paternity skew and the opportunity for selection in a mammal with reversed sexual size dimorphism. Molecular Ecology, 15, 3035-3043.


  1. Glover AM (2006) PhD Thesis (University of Leeds) The ecology and conservation of cave-roosting bats in the Yorkshire Dales.

  2. Goodship N (2006) PhD Thesis (University of Cardiff) Endocrine control of nestling begging behaviour in the pied flycatcher, Ficedula hypoleuca.

  3. Griffiths AM (2006) PhD Thesis (University of Exeter) The population structure of brown trout (Salmo salar) on Dartmoor National Park.

  4. Phillimore A (2006) PhD Thesis (Imperial College, University of London) The ecological basis of speciation and divergence in birds.

  5. Vernon C (2006) PhD Thesis (University of Liverpool) Transcriptomic assessment of gene responses to environmental stress - genetic and comparative approaches.

[NBAF-E] 3

  1. Hegarty MJ, Jones JM, Wilson ID, Barker GL, Coghill JA, Sanchez-Baracaldo P, Liu G, Buggs RJ, Abbott RJ, Edwards KJ, Hiscock SJ (2005) Development of anonymous cDNA microarrays to study changes to the Senecio floral transcriptome during hybrid speciation. Molecular Ecology, 14, 2493-2510.

  2. Jones M, Blaxter M (2005) Evolutionary biology: animal roots and shoots. Nature, 434, 1076-1077.

  3. Papanicolaou A, Joron M, McMillan WO, Blaxter ML, Jiggins CD (2005) Genomic tools and cDNA derived markers for butterflies. Molecular Ecology, 14, 2883-2897.


  1. Williams D R, Epperson LE, Li WZ, Hughes MA, Taylor R, Rogers J, Martin SL, Cossins AR, Gracey AY (2005) Seasonally hibernating phenotype assessed through transcript screening. Physiological Genomics 24, 12-22.

[NBAF-S] 15

  1. Bretman A and Tregenza T (2005) Measuring polyandry in the wild: a case study using promiscuous crickets. Molecular Ecology, 14, 2169-2179.

  2. *Carpenter PJ, Pope LC, Greig C, Dawson DA, Rogers LM, Erven K, Wilson GJ, Delahay RJ, Cheeseman CL, Burke T (2005) Mating system of the Eurasian badger, Meles meles, in a high density population. Molecular Ecology, 14, 273-284.

  3. Châline N, Martin SJ, Ratnieks FLW (2005) Absence of nepotism towards imprisoned young queens during swarming in the honey bee. Behavioral Ecology, 16, 403-409.

  4. Dawson DA, Chittock J, Jehle R, Whitlock A, Nogueira D, Pellat J, Birkhead T, Burke T (2005) Identification of 13 polymorphic loci in the zebra finch Taeniopygia guttata (Passeridae, Aves). Molecular Ecology Notes, 5, 298-301.

  5. Dawson DA, Hunter FM, Pandhal J, Buckland R, Parham A, Jones IL, Bradshaw M, Jehle R, Burke T (2005) Assessment of 17 new whiskered auklet (Aethia pygmaea) microsatellite loci in 42 seabirds identifies 5-15 polymorphic markers for each of nine Alcinae species. Molecular Ecology Notes, 5, 289-297.

  6. *Gunn MR, Dawson DA, Leviston A, Hartnup K, Davis CS, Strobeck C, Slate J, Coltman DW (2005) Isolation of 18 polymorphic microsatellite loci from the North American red squirrel, Tamiascirus ludsonicus (Sciuridae, Rodentia) and their cross-utility in other species. Molecular Ecology Notes, 5, 650-653.

  7. Leech DI, Rowe LV, Hartley IR (2005) Experimental evidence for adjustment of parental investment in relation to brood sex ratio in the blue tit. Animal Behaviour, 72,1301-1307.

  8. Pegler KR, Evans L, Stevens JR, Wall R (2005) Morphological and molecular comparison of host-derived populations of parasitic Psoroptes mites.  Medical and Veterinary Entomology, 19, 392-403. 

  9. Reichard M, Bryja J, Ondra Kova M, Davidova M, Kaniewska P, Smith C (2005) Sexual selection for male dominance reduces opportunities for female mate choice in the European bitterling (Rhodeus sericeus) Molecular Ecology, 14, 1533-1542.

  10. Rivers NM, Butlin RK, Altringham JD (2005) Genetics population structure of Natterer's bats explained by mating at swarming sites and philopatry. Molecular Ecology, 14, 4299-4312.

  11. Rossiter SJ, Ransome RD, Faulkes CG, Le Comber SL, Jones G (2005) Mate-fidelity and intra-lineage polygyny in greater horseshoe bats. Nature, 437, 408-411.

  12. Senior P, Butlin RK, Altringham JD (2005) Sex and segregation in temperate bats. Proceedings of the Royal Society, Series B, 272, 2467-2473.

  13. *Wenseleers T, Badcock N, Erven K, Tofilski A, Nascimento FS, Hart AG, Burke T, Archer ME, Ratnieks FLW (2005) A test of worker policing theory in an advanced eusocial wasp, Vespula rufa. Evolution, 59, 1306-1314.

  14. Wilcock HR, Bruford MW, Hildrew AG, Nichols RA (2005) Recruitment, kin and the spatial genetic structure of a caddisfly Plectrocnemia conspersa in a southern English stream. Freshwater Biology, 50, 1499–1514.


  1. Anderson SJE (2005) PhD Thesis (Open University) Metapopulation genetics of the garden tiger moth (Arctia caja).

  2. Barnett OE (2005) PhD Thesis (Open University) The role of environmental change in the maintenance of sexual reproduction in a perennial grass species, Festuca rubra.

  3. *Barr, I. (2005). PhD Thesis, University of Sheffield. Sexual selection in the blue tit (Parus caeruleus).

  4. Green NM (2005) PhD Thesis (University of Leeds) Seasonal patterns in population structure and behaviour of the bat Myotis nattereri.

  5. Senior P (2005) PhD Thesis (University of Leeds) Spatial and sexual segregation in Daubenton's bat (Myotis daubentonii) in Wharfedale.

  6. Temple HJ (2005) PhD Thesis (University of Cambridge) Ecology, cooperative breeding and conservation of the white-breasted thrasher Ramphocinclus brachyurus.

  7. Tingay R (2005) PhD Thesis (University of Nottingham) Historical distribution, contemporary status and cooperative breeding in the Madagascar fish eagle (Haliaeetus vociferoides): Implications for conservation.

  8. Webster L (2005) PhD Thesis (University of Glasgow) The effects of gene flow on local adaptation in a natural host-parasite system.

[2004] NBAF-S ONLY from this point on. 12

  1. Bretman A, Wedell N, Tregenza T (2004) Molecular evidence of post-copulatory inbreeding avoidance in the field cricket Gryllus bimaculatus. Proceedings of the Royal Society of London B, 271, 159-164.

  2. Châline N, Ratnieks FLW, Raine NE, Badcock NS, Burke T (2004) Non-lethal sampling of honey bee, Apis mellifera, DNA using wing tips. Apidologie, 35, 311-318.

  3. Dawson DA, Rossiter SJ, Jones G, Faulkes CG (2004) Microsatellite loci for the greater horseshoe bat Rhinolophus ferrumequinum (Rhinolophidae, Chiroptera) and their cross-utility in other 17 other bat species. Molecular Ecology Notes, 4, 96-100.

  4. Fletcher K & Hamer KC (2004) Offspring sex ratio in the common tern Sterna hirundo: a species with negligible sexual size dimorphism. Ibis, 146, 454-360.

  5. Frantz AC, Schaul M, Pope LC, Fack F, Schley L, Muller CP, Roper TJ (2004) Estimating population size by genotyping remotely plucked hair: the Eurasian badger. Journal of Applied Ecology, 41, 985-995.

  6. Hatchwell BJ, Russell AF, MacColl ADC, Ross DJ, Fowlie MK, McGowan A (2004) Helpers increase long-term but not short-term productivity in cooperatively breeding long-tailed tits. Behavioral Ecology, 15, 1-10.

  7. Hay JM, Evans PR, Ward RM, Hamer KC (2004) Poor nutritional condition as a consequence of high dominance status in the coal tit, Parus ater. Ibis, 146, 103-107.

  8. Kingston T, Rossiter S (2004) Harmonic-hopping in Wallacea's bats. Nature, 429, 654-657.

  9. Madden JR, Endler JA, Jury F (2004) Morphological signals of sex and status in spotted bowerbirds. Emu, 104, 21-30.

  10. Madden JR, Lowe TJ, Fuller HV, Coe RL, Dasmahapatra KK, Amos W, Jury F (2004) Neighbouring male spotted bowerbirds are not related, but do maraud each other. Animal Behaviour, 68, 751-758.

  11. Madden JR, Lowe TJ, Fuller HV, Dasmahapatra KK, Coe RL (2004) Local traditions of bower decoration by spotted bowerbirds in a single population. Animal Behaviour, 68, 759-765.

  12. Nichols RA, Freeman KLM (2004) Using molecular markers with high mutation rates to obtain estimates that distinguish relative migration rate and population size: a demonstration using data from endemic Mauritian skinks. Molecular Ecology, 13, 775-789.


  1. Bretman A.J. (2004) PhD Thesis (University of Leeds) Sexual selection in the field cricket Gryllus bimaculatus.

  2. Chaline N.G. (2004) PhD Thesis (University of Sheffield) Reproductive conflicts and nepotism in the honey bee.

  3. Evans L.M. (2004) PhD Thesis (University of Exeter) Phylogenetic assessment of Psoroptes ovis; their epidemiology and taxonomic identity explored.

  4. Frantz A.C. (2004) PhD Thesis (University of Sussex) Non-invasive genetic typing in the study of badger (Meles meles) ecology.

  5. *Furmankiewicz J. (2004) PhD Thesis (University of Wroclaw, Poland. [in Polish] with time spent in Leeds)
    Mating behaviour of brown long-eared bat Plecotus auritus (Linnaeus, 1758).

  6. *Paszkiewicz R. (2004) PhD Thesis (Wroclaw University, Poland - with time spent at Sunderland University) Genetical structure and population biology of the mouse-eared bat (Myotis myotis).

  7. Pedro PM (2004) PhD Thesis (University of Leeds) Impact of habitat fragmentation on species of Sabethes mosquitoes.

[2003] 16

  1. *Berg PR, Dawson DA, Pandhal J, Kirdendall LR, Burke T (2003) Isolation of microsatellite loci from two inbreeding bark beetle species (Coccotrypes). Molecular Ecology Notes, 3, 270-273.

  2. *Carpenter PJ, Dawson DA, Greig C, Parham A, Cheeseman CL, Burke T (2003) Isolation of 39 polymorphic microsatellite loci and the development of a fluorescent marker set for the Eurasian badger (Meles meles) (Carnivora: Mustelidae). Molecular Ecology Notes, 3, 610-615.

  3. Davies NB, Butchart SHM, Burke TA, Châline N, Stewart IR (2003) Reed warblers guard against cuckoos and cuckoldry. Animal Behaviour, 65, 285-295.

  4. Dawson DA, Bretman AJ, Tregenza T (2003) Microsatellite loci for the field cricket, Gryllus bimaculatus and their cross-utility in other species of Orthoptera. Molecular Ecology Notes, 3, 191-195.

  5. Dawson DA, Burland TM, Douglas A, LeComber SC, Bradshaw M (2003) Isolation of microsatellite loci in the freshwater fish, the bitterling Rhodeus sericeus (Teleostei: Cyprinidae). Molecular Ecology Notes, 3, 199-202.

  6. Evans LM, Dawson DA, Wall R, Burke T, Stevens J (2003) Isolation of Psoroptes scab mite microsatellite markers (Acari: Psoroptidae). Molecular Ecology Notes, 3, 420-424.

  7. Fletcher KL & Hamer KC (2003) Sexing terns using biometrics: the advantage of within-pair comparisons. Bird Study, 50, 78-83.

  8. Frantz AC, Pope LC, Carpenter PJ, Roper TJ, Wilson GJ, Delahay RJ, Burke T (2003) Reliable microsatellite genotyping of the Eurasian badger (Meles meles) using faecal DNA. Molecular Ecology, 12, 1649-1661.

  9. Frentui F, Lange CL, Burke T, Owens IPF (2003) Isolation of microsatellite loci in the Capricorn silvereye, Zosterops lateralis chlorocephalus (Aves: Zosteropidae). Molecular Ecology Notes, 3, 462-465.

  10. *Jump AS, Woodward FI, Burke T (2003) Cirsium species show disparity in patterns of genetic variation at their range-edge, despite similar patterns of reproduction and isolation. New Phytologist, 160, 359-370.

  11. Madden JR (2003) Bower decorations are good predictors of mating success in the spotted bowerbird. Behavioural Ecology and Sociobiology, 53, 269-277.

  12. Madden JR (2003) Male spotted bowerbirds preferentially choose, arrange and proffer objects that are good predictors of mating success. Behavioural Ecology and Sociobiology, 53, 263-268.

  13. Marshall R, Buchanan KL, Catchpole CK (2003) Sexual selection and individual genetic diversity in a songbird. Proceedings of the Royal Society of London, Series B (suppl.), S248-S250.

  14. McGowan A, Woodburn RJW, Hatchwell BJ (2003) The effect of helping behaviour on the survival of juvenile and adult long-tailed tits (Aegithalos caudatus). Journal of Animal Ecology, 72, 491-499.

  15. *Wilhelm K, Dawson DA, Gentle LK, Horsfield GF, Schlötterer C, Greig C, East M, Hofer H, Tautz D, Burke T (2003) Characterisation of spotted hyena, Crocuta crocuta microsatellite loci. Molecular Ecology Notes, 3, 360-362.

  16. Wilson GJ, Frantz AC, Pope LC, Roper TJ, Burke TA, Cheeseman CL, Delahay RJ (2003) Estimation of badger abundance using faecal DNA typing. Journal of Applied Ecology, 40, 658-666.


  1. Ahmed S. (2003) PhD Thesis (University of Leeds) Pollen mediated gene flow in Ficus sycomorus L.

  2. Douglas A. (2003) PhD Thesis (Queen Mary, University of London) Population ecology and behaviour of juvenile bitterling (Rhodeus serecius, Pallas).

  3. Freeman K. (2003) PhD Thesis (Queen Mary, University of London) The population genetics and conservation of a Mauritian skink.

  4. Hay J. (2003) PhD Thesis (University of Durham) Causes and consequences of variation in dominance status in the Paridae.

  5. Sharp S.P. (2003) PhD Thesis (University of Sheffield) Kin recognition in the cooperative breeding system of long-tailed tits Aegithalos caudatus.

[2002] 9

  1. Châline N, Ratnieks FLW, Burke T (2002) Anarchy in the UK: Detailed analysis of worker reproduction in a naturally-occurring British anarchistic honeybee, Apis mellifera, colony using microsatellite markers. Molecular Ecology, 11, 1795-1803.

  2. Dawson DA & Wilcock HR (2002) Isolation and characterisation of eight polymorphic microsatellite loci in the net-spinning caddisfly, Polycentropus flavomaculatus (Polycentropodidae). Molecular Ecology Notes, 2, 514-517.

  3. Hatchwell BJ, Ross DJ, Châline N, Fowlie MK, Burke T (2002) Parentage in the cooperative breeding system of long-tailed tits, Aegithalos caudatus. Animal Behaviour, 64, 55-63.

  4. *Jump AS, Dawson DA, James CM, Woodward FI, Burke T (2002) Isolation of polymorphic microsatellites in the stemless thistle (Cirsium acaule) and their utility in other Cirsium species. Molecular Ecology Notes, 2, 589-592.

  5. *Krupa AP, Jehle R, Dawson DA, Gentle LK, Gibbs M, Arntzen JW, Burke T (2002) Microsatellite loci in the crested newt (Triturus cristatus) and their utility in other newt taxa. Conservation Genetics, 3, 87-89.

  6. MacColl ADC & Hatchwell BJ (2002) Temporal variation in fitness pay-offs promotes cooperative breeding in long-tailed tits Aegithalos caudatus. American Naturalist, 160, 186-194.

  7. Madden JR (2002) Bower decorations attract females but provoke other male spotted bowerbirds - bower owners resolve this trade off. Proceedings of the Royal Society, Series B, 269, 1347-1351.

  8. Phillips RA (2002) Trios of brown skuas at Bird Island, South Georgia: incidence and composition. Condor, 104, 694-697.

  9. Phillips RA, Dawson DA, Ross DJ (2002) Mating patterns and reversed size dimorphism in Southern skuas (Stercorarius skua lonnbergi). Auk, 119, 858-863.
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