Comments: (1) The following genera and species proposed by Tzvelev are omitted from the edited draft as represented only by casuals in the Arctic




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37  Poaceae (Draft: N.N. Tzvelev)

S Gramineae

Comments:

(1) The following genera and species proposed by Tzvelev are omitted from the edited draft as represented only by casuals in the Arctic   Anisantha C. Koch (A. tectorum (L.) Nevski), Apera Adans. (A. spica-venti (L.) Beauv.), Avena L. (A. sativa L.), Bromus L. s. str. (B. arvensis L., B. hordeaceus L., B. secalinus L.), Ceratochloa DC. & Beauv. (C. marginata (Nees. ex Steud.) B.D. Jacks.), Polypogon Desf. (P. monspeliensis (L.) Desf.), Secale L. (S. cereale L.) and Triticum L. (T. aestivum L.)   or as not present at all   Holcus L. (H. lanatus L.). Omitted genera and species are listed at the end. One genus   Pseudoroegneria   is added to Tzvelev's draft. (Elven)



37.1  Elymus L. (1753), Sp. Pl. 83.

S Agropyron Gaertn., p.p.; Roegneria C. Koch (1848), Linnaea 21: 413.

Comments:

(1) Three sections: sect. Elymus (E. confusus), sect. Hystrix (Moench) Á. Löve (E. asiaticus), and sect. Goulardia (Husnot) Tzvelev (all the others).

Two of the three sections given in Tzvelev's draft contain only one species each. Sections are therefore omitted in this edition. They could be entered (as the genus has more than 10 species), or they could better be replaced by an initial comment like the one above. (Elven)

(2) Species delimitation and names have been in a flux in this genus for a long time, and still is. This is a serious hindrance for all users of botanical nomenclature, not only taxonomists but also phytogeographers, ecologists and conservationists. As the checklist also aims to serve the non-taxonomists we should try not to introduce too many novelties compared with previous accounts, novelties that might not survive the checklist phase.

In his draft, Tzvelev proposes new solutions in many cases compared with both his previous treatments and the previous treatments of others. The number of species proposed by Tzvelev is not excessive, but the subspecific split might be so, and the names proposed at species level are in several cases stange compared with the previous treatments of him and others. I think we need to see very good justifications, based on morphology and/or genetics, before we introduce even more complications in Elymus (and Elytrigia).

However, except for the separation of Pseudoroegneria, the edited draft is almost identical to Tzvelev's proposal but I have given my views in many small comments. I think we need to ask a small group, combining expertise from the Russian and American side (Mary Barkworth?) and perhaps from the 'genome' group in Sweden (Björn Salomon?), if they are willing to work through this genus before a final checklist proposal is reached. (Elven)


37.1.1  Elymus mutabilis (Drobov) Tzvelev in Grubov (1968), Pl. As. Centr. 4: 217.

B Agropyron mutabile Drobov (1916), Trudy Bot. Muz. Imp. Akad. Nauk 16: 88.

S Roegneria mutabilis (Drobov) Hyl. (1945), Ups. Univ. Årsskr. 7: 36.

T Siberia: Yakutia, bank of Chona River near village Dushenki, 1914, leg. Drobov 315 (LE), lectotype selected by ***.

2n= 28 (4x).

2nD Löve & Löve (1975) list several counts, one as arctic.

G RUS SIB

Comments:

(1) Approaches, but does not reach, the Arctic also in N Norway. (Elven)
The Elymus trachycaulus aggregate (E. subsecundus, E. trachycaulus)

Comments:

(1) A large, intricate and mainly North American -Greenlandic complex where the 'subsecundus' entity might be included as a subspecies in a more widely defined E. trachycaulus, or where several species might be recognized. The current treatment reflects the imbalance between North American and Russian approaches; the former recognize subspecies (or rather varieties) where the latter recognize species. The American taxa listed below as subspecies of E. trachycaulus differ in more characters, and are less sympatric, than many of the entities listed as species in the next aggregate (E. violaceus agg.). (Elven)
37.1.2  Elymus subsecundus (Link) Á. Löve & D. Löve (1964), Taxon 13: 201.

B Triticum subsecundum Link (1833), Hort. Bot. Berol. 2: 190.

S Agropyron subsecundum (Link) Hitchc. (1934), Amer. J. Bot. 21: 131; Elymus trachycaulus (Link) Gould ex Shinners subsp. subsecundus (Link) Á. Löve & D. Löve (1976), Bot. Not. 128: 502; Agropyron trachycaulon (Link) Malte ex H.F. Lelois [Lewis?] var. unilaterale (Cassidy) Malte (1932), Ann. Rep. Natl. Mus. Can. 1930: 42.

2n= 28 (4x).

2nD Löve & Löve (1975) list several non-arctic counts.

G ALA


Comments:

(1) To be treated as separate species or as part of an extended E. trachycaulus complex? The relevant subspecies combination is available. (Elven)

(2) Mapped by Hultén (1968) from the Kotzebue area in Alaska. Therefore a border case if it has not been found in new sites. (Elven)

WARNING! Will be reduced to a subspecies of E. trachycaulus if not well justified.
37.1.3  Elymus trachycaulus (Link) Gould ex Shinners (1954), Rhodora 56: 28.

B Triticum trachycaulon Link (1833), Hort. Bot. Berol. 2: 189.

S Agropyron trachycaulon (Link) Malte ex H.F. Lelois [Lewis??] (1931), Can. Field Natural. 45: 201.

Comments:

(1) Hultén (1968, Ark. Bot., ser. 2, 7, 1: 20) argues that Triticum pauciflorum Schwein. in Keat. (1824), Narr. Exped. St. Peter's River 2: 383, is the oldest name at species level in the group. It is even, acc. to him, valid within Agropyron as A. pauciflorum Schur (1859), Verh. Siebenb. Ver. Naturw. 10: 77, is invalid (published in synonymy of A. caninum). However, there might be an earlier Elymus pauciflorus I don't know about. Opinions? (Elven)
37.1.3.1  Elymus trachycaulus (Link) Gould ex Shinners subsp. trachycaulus

S Agropyron trachycaulon (Link) Malte ex H.F. Lelois [Lewis?] var. trachycaulon; Agropyron pauciflorum (Schwein.) Hitchc. subsp. pauciflorum.

2n= 28 (4x)

2nD ?


G RFE ALA CAN

Comments:


37.1.3.2  Elymus trachycaulus (Link) Gould ex Shinners subsp. major (Vasey) Tzvelev (1973), Nov. Sist. Vyssh. Rast. 10: 24.

B Agropyron violaceum (Hornem.) Lange var. majus Vasey (1893), Contrib. U.S. Natl. Herb. 1: 280.

S Agropyron pauciflorum (Schwein.) Hitchc. subsp. majus (Vasey) Melderis in Hultén (1968), Ark. Bot., ser. 2, 7, 1: 20.

2n= 28 (4x).

2nD Schulz-Schaeffer & Jurasits (1967 **).

G ALA


Comments:

(1) Mapped by Hultén (1968) from one 'dot' near Nome in Seward Peninsula. Confirmed from here? (Elven)


37.1.3.3  Elymus trachycaulus (Link) Gould ex Shinners subsp. novae-angliae (Scribn.) Tzvelev (1973, Nov. Sist. Vyssh. Rast. 10. 23

B Agropyron tenerum (Malte) *** var. novae-angliae Scribn. in Farwell (1920), Rep. Mich. Acad. Sci. 21: 355.

S Agropyron trachycaulum (Link) Malte ex H.F. Lelois [Lewis?] var. novae-angliae (Scribn.) Fern. (1933), Rhodora 35: 174; Agropyron pauciflorum (Schwein.) Hitchc. subsp. novae-angliae (Scribn.) Melderis in Hultén (1968), Ark. Bot., ser. 2, 7, 1: 20.

2n=


2nD

G RFE


Comments:

(1) Absent from Tzvelev's draft but reported by Tzvelev (1976: 119) for "Far East: Arctic (near Senyavinskie hot water springs, perhaps wild)". As this locality is in Chukotka Peninsula, it is within the Arctic as defined. (Tzvelev & Elven)


37.1.3.4  Elymus trachycaulus (Link) Gould ex Shinners subsp. virescens (Lange) Á. Löve & D. Löve (1976), Bot. Not. 128: 502.

B Agropyron violaceum (Hornem.) Lange var. virescens Lange (1880), Medd. Grønl. 3: 155.

S

2n= 28 (4x).



2nD Löve & Löve (1975) list four counts, two Greenlandic.

G ALA? GRL

Comments:

(1) The occurrence of this Greenland taxon in Alaska needs careful documentation. I am not aware of any reliable report of it from there. Alaska will be excluded if not documented. (Elven)


The Elymus violaceus aggregate (E. alaskanus, E. kronokensis, E. sajanensis, E. violaceus)

Comments:

(1) This is the second aggregate where most entities at one time or the other have been recombined under a majority of the names. In his current draft and partly in 1973, Tzvelev proposes recombination of the 'andinus' entity under E. violaceus (previously under E. trachycaulus), the 'hyperarcticus' entity under E. sajanensis (previously under E. alaskanus or E. violaceus), the 'villosus' entity under E. sajanensis (previously under E. alaskanus), the 'borealis' entity under E. kronokensis (previously as separate or under E. alaskanus), and the 'subalpinus' entity under E. kronokensis (previously as separate or under E. alaskanus). It seems to be extremely difficult to agree on where the limits between species should run.

The impression received by an ignorant reader of Elymus (like me) is that we might solve some of the problems by rising the species limit one step. We could consider some of the species accepted in the draft as subspecies, and some of the subspecies as locally deviating population groups (e.g., 'villosus', 'caeruleus', and 'scandicus'). (Elven)


37.1.4  Elymus violaceus (Hornem.) Feilberg (1984), Medd. Grønl., Biosci. 15: 12.

B Triticum violaceum Hornem. (1832), Fl. Dan. [f. 35], t. 2044.

S Agropyron violaceum (Hornem.) Lange (1880), Medd. Grønl. 3: 155; E. trachycaulus (Link) Gould ex Shinners subsp. violaceus (Hornem.) Á. Löve & D. Löve (1976), Bot. Not. 128: 502.

Comments:

(1) To be treated as a separate species or as part of an extended E. trachycaulus complex? The Cordilleran 'andinus' entity has, e.g., fairly constantly been treated as part of E. trachycaulus. (Elven)

(2) The combination Agropyron violaceum seems to have been published earlier by Lange in Rink (1877), Greenland 115. This is also the citation of Lange (1880). (Elven)


37.1.4.1  Elymus violaceus (Hornem.) Feilberg subsp. violaceus

S Agropyron violaceum (Hornem.) Lange subsp. violaceum.

2n= 28 (4x).

2nD Löve & Löve (1975) list two Greenlandic counts.

G GRL

Comments:


37.1.4.2  Elymus violaceus (Hornem.) Feilberg subsp. latiglumis (Scribn. & J.G. Sm.) Tzvelev (2000), Nov. Sist. Vyssh. Rast. 32: 181.

B Agropyron violaceum (Hornem.) Lange var. latiglume Scribn. & J.G. Sm. (1897), Bull. U.S. Dept. Agric. Div. Agrost. 4: 30.

S

T USA: Montana, Gallatin Co., Lone Mountain, leg. Twedy 1011 (US) lectotype, see Hitchcock & Chase (1951: 798).



2n= 28 (4x).

2nD ?


G ALA

Comments:


37.1.4.3  Elymus violaceus (Hornem.) Feilberg subsp. andinus (Scribn. & J.G. Sm.) Tzvelev (2000), Nov. Sist. Vyssh. Rast. 32: 182.

B Agropyron violaceum (Hornem.) Lange var. andinum Scribn. & J.G. Sm. (1897), Bull. U.S. Dept. Agric. Div. Agrost. 4: 30.

S Agropyron violaceum (Hornem.) Lange subsp. andinum (Scribn. & J.G. Sm.) Melderis in Hultén (1968), Ark. Bot., ser. 2, 7, 1: 19; Elymus trachycaulus (Link) Gould ex Shinners subsp. andinus (Scribn. & J.G. Sm.) Á. Löve & D. Löve (1976), Bot. Not. 128: 502.

2n= 28 (4x).

2nD ?

G ALA


Comments: See comment under the species.
37.1.5  Elymus alaskanus (Scribn. & Merr.) Á. Löve (1970), Taxon 19: 299.

B Agropyron alaskanum Scribn. & Merr. (1910), Contr. U.S. Natl. Herb. 13: 85.

S E. alaskanus (Scribn. & Merr.) Á. Löve subsp. alaskanus; Agropyron boreale (Turcz.) Drobov subsp. alaskanum (Scribn. & Merr.) Melderis in Hultén (1968), Ark. Bot., ser. 2, 7, 1: 19.

T Alaska: Circle City, 18.08.1899, leg. W.H. Osgood (US) holotype.

2n= 28 (4x).

2nD Hedberg (1967 Can?); Knaben (1968 Ala).

G RFE ALA CAN

Comments:



(1) How the 'alaskanus' entity is treated is crucial to the nomenclature of this and the next species in Tzvelev's draft as Agropyron alaskanum has priority (1910) before Agropyron kronokense (1915), and also before Roegneria sajanensis (1934). Hultén (1943, Lunds Univ Årsskr., n. f., avd. 2, 38, 1: 256) writes about Agropyron alaskanum: "The peculiar glumes as well as the pilose nodes seem. however, to unite all these different forms into a fairly distinct type. The hairiness of the nodes is very variable, sometimes rather scanty but sometimes almost covering the surface...". I am not at all sure this is enough to justify E. alaskanus as a species separate from the others in the aggregate. This was also the view of Melderis in Tutin et al. (1980). He considered E. kronokensis a synonym of E. alaskanus and united the entities named as 'alaskanus', 'borealis', 'scandicus' and 'subalpinus' inside this species for the European parts. (Elven)


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