86 Boraginaceae




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86  Boraginaceae (Draft: V.V. Petrovsky)

Comments:

(1) Three genera in Petrovsky's draft are excluded (Amsinckia, Cryptantha and Lycopsis) and two more are in danger (Asperugo, Lappula).

Amsinckia Lehm. (1831), Sem. Hort. Bot. Hamburg. 1: 7, with two species. Both are most probably casual in the Arctic and should be excluded: A. menziesii (Lehm.) Nelson & Macbride (1916), Bot. Gaz. (Crawfordsville) 61: 36 [basionym: Echium menziesii Lehm. (1830), Nov. Stirp. Pug. 2: 29]   given from ALA; and A. lycopsoides (Lehm.) Lehm. (1831), Sem. Hort. Bot. Hamburg. 1831: 3 [basionym: Lithospermum lycopsoides Lehm. (1830), Nov. Stirp. Pug. 2: 28]   also given from ALA. (Elven)

Cryptantha Lehm. ex Fisch. & Mey. (1836), Ind. Sem. Hort. Petrop. 2: 35, with C. spiculifera (Piper) Payson (1927), Ann. Miss. Bot. Gard. 14: 298 [basionym: Oreocarya spiculifera Piper (1906), Contr. U.S. Nat. Herb. 11: 481]. Reported from RFE. The Chukotka record is based on a single casual occurrence, but probably as a native (or non-anthropogenic) casual. The identity of the Chukotka plant with this species is doubted by D.F. Murray. This only Asian specimen of this American genus should be checked by some American expert before acceptance.

Lycopsis L. (1753), Sp. Pl. 138, with L. arvensis L. (1753), Sp. Pl. 139 [synonym: Anchusa arvensis (L.) M. Bieb. (1808), Fl. Taur. Caucas. 1: 123]. Given with 2n=48 (8x); Lövkvist (1963 Sweden); Gadella & Kliphuis (1970 S Eur). Reported from N Fennoscandia as a casual weed. It is obviously a much too thermophilous species to be stable anywhere in the Arctic. Therefore omitted, and then we need not decide between Lycopsis or Anchusa.

For Asperugo and Lappula, see comments below. (Elven)



86.1  Eritrichium Schrad. (1820), Comm. Gotting. 4: 186, in obs.

Comments:

(1) Löve & Löve (1975) list another species, E. czekanowskii Trautv. (1877), Acta Hort. Petrop. 5: 88, from Siberia. The name is absent from Fl. Arct. URSS 8. Is it a worthwhile taxon, and if so does it occur within (or near) the Arctic? (Elven)
86.1.1  Eritrichium villosum (Ledeb.) Bunge (1836), Verz. Pfl. Altai Gebirge 14.

B Myosotis villosa Ledeb. (1815), Mém. Acad. Sci. Pétersb. 5: 516.

S

2n= For the collective species. (1) 24 (4x, x=6). (2) 48 (8x, x=6).



2nD (1) Zhukova (1968, 1969); Zhukova & Tikhonova (1973); Zhukova et al. (1973). (2) Zhukova et al. (1973).

Comments:

(1) The chromosome counts were published before the recognition of subsp. pulvinatum. Petrovsky should try to sort the numbers on subspecies. (Elven)

(2) The two subspecies are nearly sympatric   subsp. pulvinatum occurs within the range of subsp. villosum. How different are they and what keeps them apart? (Elven)



WARNING! Subspecies might be omitted if not sufficiently well differentiated.
86.1.1.1  Eritrichium villosum (Ledeb.) Bunge subsp. villosum

S

2n=



2nD

G RUS SIB RFE

Comments:
86.1.1.2  Eritrichium villosum (Ledeb.) Bunge subsp. pulvinatum Petrovsky in Tolm. & Jurtz. (1980), Fl. Arct. URSS 8: 241.

S

T Type: Siberia: Pars orientalis peninsulae Taimyr, sinus Mariae Pronczisczeviae, 26.07.1972, leg. N. Matveeva 380 (LE) holotypus.



2n=

2nD


G SIB RFE

Comments:


86.1.2  Eritrichium sericeum (Lehm.) DC. (1846), Prodr. 10: 126.

B Myosotis sericea Lehm. (1818), Pl. Asperif. Nucif. 2: 98.

S

2n= For the collective species. 28 (4x, x=7).



2nD Zhukova & Petrovsky (1971); Zhukova et al. (1973).

Comments:

(1) The chromosome reports counts were published before the recognition of subsp. arctisibiricum. Petrovsky should try to find out to which of the subspecies they belong. (Elven)

(2) The distribution of subsp. arctisibiricum is very strange, disjunct from Taimyr with surroundings to Wrangel Island whereas the taxon between these areas is subsp. sericeum. This is not a consistent phytogeographic pattern and needs some explanation before the taxa are fully accepted as significantly different. (Elven)



WARNING! Subspecies might be omitted if not justified as sufficiently differentiated.
86.1.2.1  Eritrichium sericeum (Lehm.) DC. subsp. sericeum

S

2n=



2nD

G SIB RFE

Comments:
86.1.2.2  Eritrichium sericeum (Lehm.) DC. subsp. arctisibiricum Petrovsky in Tolm. & Jurtz. (1980), Fl. Arct. URSS 8: 240.

S

T Type: Russian Far East: Insula Wrangelii (pars occidentalis) in cursu medio fl. Gusinaja, in valli rivuli Korall, 02.07.1970, leg. P. Zhukova & V.V. Petrovsky 70-48 (LE) holotype.



2n= 28 (4x, x=7).

2nD Petrovsky in Tolmachev & Yurtsev (1980 NE As).

G SIB RFE

Comments:


86.1.3  Eritrichium splendens Kearney ex W.F. Wight (1902), Bull. Torrey Bot. Club 1920: 410.

S

T Alaska: Old Man Creek, 09.07.1901, leg. Mendenhall (US) holotype.



2n= 36 (6x, x=6).

2nD Johnson & Packer (1968 Ala).

G ALA

Comments:


86.1.4  Eritrichium aretioides (Cham.) DC. (1846), Prodr. 10: 125.

B Myosotis aretioides Cham. (1829), Linnaea 4: 443.

S

T [Described from the Bering Strait area: "Ad sinum St Laurentii ... in insula Sancti Laurentii", probably erroneous in St. Lawrence Island.]



Comments:
86.1.4.1  Eritrichium aretioides (Cham.) DC. var. aretioides

S

2n= 24 (4x, x=6).



2nD Löve & Löve in Löve (1975d).

G RFE ALA

Comments:

(1) This variety seems to be excluded from the arctic in Fl. Arct. URSS 8 (p. 243). (Elven)


86.1.4.2  Eritrichium aretioides (Cham.) DC. var. chamissonis (DC.) Petrovsky in Tolm. & Jurtz. (1980), Fl. Arct. URSS 8: 243.

B E. chamissonis DC. (1846), Prodr. 10: 125.

S

T [Described from Russian Far East: "Ad Sinum Sancti Laurentii ad fretum Beering ... et Kamtchatka".]



2n= 48 (8x, x=6).

2nD Johnson & Packer (1968 Ala); Zhukova et al. (1973 NE As).

G RFE ALA

Comments:


86.1.5  Eritrichium tschuktschorum Jutz. & Petrovsky in Tolm. Jurtz. (1980), Fl. Arct. URSS 8: 244.

S

T Type: Russian Far East: Districtus autonomicus Tschukotskij, planiteis elata Anjuensis, in systemate fl. Malyi Anjui, ad fontes fl. Koralveem, prope opidulum Bezymyannyi, 23.06.1966, leg. P. Zhukova & V.V. Petrovsky 66-44 (LE) holotype.



2n= 20 (x=?).

2nD Petrovsky in Tolmachev (1980 NE As).

G RFE

Comments:



86.2  Mertensia Roth (1797), Catal. Bot. 1: 34.
86.2.1  Mertensia maritima (L.) Gray (1821), Nat. Arr. Brit. Pl. 2: 354.

B Pulmonaria maritima L. (1753), Sp. Pl. 136.

S

T [Described from Britain.]



2n= 24 (4x).

2nD Löve & Löve (1975) list several counts for the species, most as arctic.

Comments:

(1) The chromosome counts listed by Löve & Löve (1975) probably include both varieties. (Elven)

(2) The arctic race   var. tenella   seems to be both morphologically and genetically distinct from the more southern var. maritima. Skarpås (1999) found total identity in genetical markers (enzymes) in var. maritima from southernmost to northernmost mainland Norway and clear differences from arctic (Svalbard) material og var. tenella. Var. tenella is described from Svalbard. The arctic race is also proved (morphologically) from Jan Mayen, Greenland and arctic Canada whereas, e.g., all the Icelandic material I have seen is var. maritima.

The identity of the north Alaskan and east Russian material should also be checked. The plant illustrated in Hultén (1968) is a typical var. tenella and the drawing is made from an Alaskan plant. The west Russian (Kola) material is var. maritima. The arctic race probably deserves status as subspecies but does not seem to have been described as such yet. (Elven)



WARNING! The varieties should probably be raised to rank of subspecies.
86.2.1.1  Mertensia maritima (L.) Gray var. maritima

S

2n= 24 (4x).



2nD Löve & Löve (1956 Icel).

G ICE NOR RUS RFE? ALA? CAN?

Comments:

(1) The question marks for RFE, ALA and CAN refer to which race; the species is present in all areas. (Elven)


86.2.1.2  Mertensia maritima (L.) Gray var. tenella Th. Fr. (1869), Öfvers. K. Vet.-Akad. Förh. 26: ***

S

T [Described from Svalbard.]



2n=

2nD


G NOR RFE? ALA? CAN GRL

Comments:

(1) The question marks for RFE, ALA and CAN refer to which race. (Elven)
86.2.2  Mertensia drummondii (Lehm.) G. Don (1838), Gen. Hist. 4: 319.

B Lithospermum drummondii Lehm. (1830), Nov. Stirp. Pug. 2: 26.

S

2n=


2nD

G ALA CAN

Comments:
86.2.3  Mertensia sibirica (L.) G. Don (1838), Syst. Veg. 4: 319.

B Pulmonaria sibirica L. (1753), Sp. Pl. 135.

S

2n=


2nD

G SIB


Comments:
86.2.4  Mertensia pubescens (Willd. ex Roem. & Schult.) DC. (1846), Prodr. 10: 90.

B Pulmonaria pubescens Willd. ex Roem. & Schult. (1819), Syst. Veg. 4: 744.

S M. kamczatica (Turcz.) DC. (1846), Prodr. 10: 100.

2n=


2nD

G RFE


Comments:

WARNING! Might be reduced to a race of M. paniculata.
86.2.5  Mertensia paniculata (Ait.) G. Don (1838), Gen. Hist. 4: 318.

B Pulmonaria paniculata Ait. (1789), Hort. Kew., ed. 1, 1: 181.

S

2n= For the collective species. 24 (4x).



2nD Löve & Löve in Löve (1975d).

Comments:

(1) Three varieties are accepted in American standard sources and must either be accepted as races or included in the synonymy. Two of these are var. paniculata and var. alaskana (Hultén 1968, Porsild & Cody 1980, Cody 1996). The third, var. eastwoodae, is accepted as a species in Petrovsky's draft but should be justified as such. Due to Hultén's arguments (Hultén 1968, Ark. Bot.) I have reduced it to a variety in this draft version. Mertensia pubescens belongs in the same complex and should be re-evaluated with the others. (Elven)
86.2.5.1  Mertensia paniculata (Ait.) G. Don var. paniculata

S

2n=



2nD

G ALA CAN

Comments:
86.2.5.2  Mertensia paniculata (Ait.) G. Don var. alaskana (Britt.) O.L. Williams (1937), Ann. Miss. Bot. Gard. 24: 48.

B Mertensia alaskana Britt. (1901), Bull. New York Bot. Gard. 2, 6: 181.

S

T [Described from Alaska.]



2n=

2nD


G ALA? CAN

Comments:

(1) A border case in the Porcupine River valley and Richardson Mts in Yukon Territory, possibly also on the Alaskan side. (Elven)
86.2.5.3  Mertensia paniculata (Ait.) G. Don var. eastwoodae (Macbr.) Hultén (1968), Ark. Bot., ser. 2, 7, 1: 116.
B Based on M. alaskana Eastwood (1902) non Britton (1901).

S M. eastwoodae Macbride (1917), Contrib. Gray Herb., n. s., 49: 18.

T [Described from Alaska: Nome.]

2n=


2nD

G ALA


Comments:


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