15 Typhaceae (Draft: N. N. Tzvelev) s sparganiaceae (incl.) Comments: (1) Tzvelev proposes to include Sparganiaceae in Typhaceae. This is also the solution chosen by Kubitzki




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15  Typhaceae (Draft: N.N. Tzvelev)

S Sparganiaceae (incl.)

Comments:

(1) Tzvelev proposes to include Sparganiaceae in Typhaceae. This is also the solution chosen by Kubitzki (1998) but not by Kaul (2000). The obvious alternative is to retain Sparganiaceae as a family, as done by Kaul. Opinions? Tzvelev's subfamily Sparganioideae Sakht. is omitted as only one genus reaches the Arctic. (Elven)



15.1  Sparganium L. (1753), Sp. Pl. 971.

Comments:

(1) With only 5-6 species, Tzvelev's proposed division on two sections is omitted: Natantia Aschers. & Graebn. with species 1-4, Minima Aschers. & Graebn. with species 5-6. The order is also changed to place the closely related S. emersum and S. multipedunculatum together and they are emphasized as an aggregate. (Elven)
The Sparganium angustifolium-emersum aggregate (S. angustifolium, S. emersum, S. multipedunculatum)

Comments:

(1) The past and current West European and Russian view is that S. angustifolium and S. emersum (previously S. simplex) are two clearly different and perhaps not very closely related species. The most current North American view (Kaul 2000, Fl. N. Amer. 22) is rather different: "Sparganium emersum is easily confused with S. angustifolium ..." and "... the entire S. emersum-S. angustifolium complex, which is perhaps better treated as one variable speciesm S. angustifolium ...". One reason for this discrepancy is the difference in treatments of the 'multipedunculata' entity.

Hultén, a European by origin, discussed S. multipedunculata versus S. simplex/emersum, never versus S. angustifolium. The criteria he gave (Hultén 1962, Circump. Pl. 1) were that S. multipedunculatum has shorter and thicker stigmas than S. emersum and more approximate staminate heads. These quantitative features might perhaps justify subspecies rather than species but I don't know whether a subspecific combination has been made.

Kaul (2000) includes the 'multipedunculata' names in the synonymy og S. angustifolium, without any indication that they at least up to the 1970ies mostly were related to a taxon of the S. emersum/simplex group. It might be that the 'multipedunculata' entity   if there is an entity   connects S. angustifolium and S. emersum in N America whereas these two latter are more distant in Eurasia. At least, from the past history and synonymy we cannot accept a reduction of the 'multipedunculata' entity to full synonymy of S. angustifolium without some further evaluation. I have therefore very tentatively kept all three as 'species' in this draft. (Elven)
15.1.1  Sparganium emersum Rehm. (1871), Verh. Naturf. Vereins Brünn 10, Abr. 80.

S S. simplex Hudson (1778), nom. illegit., Fl. Angl., ed. 2, 401.

2n= 30 (2x).

2nD Löve & Löve (1975) list several non-arctic counts.

G RUS SIB

Comments:

(1) Kent gives 1870 as the year of publication of the name S. emersum, Kaul gives 1872. Check. (Elven)
15.1.2  Sparganium multipedunculatum (Morong) Rydb. (1905), Bull. Torrey Bot. Club 1905: 598.

B S. simplex Hudson var. multipedunculatum Morong (1888), Bull. Torrey Bot. Club 15: 79.

S

2n= 30 (2x).



2nD Löve (1954b Vegetatio 5-6: 212-224); Löve & Löve in Löve (1975b).

G ALA CAN

Comments:
15.1.3  Sparganium angustifolium Michx. (1803), Fl. Bor.-Amer. 2: 189.

S S. affine Schnizl. (1845), Typhac. 27.

2n= 30 (2x).

2nD Löve & Löve (1975) list four counts, one Icelandic.

G ICE NOR RUS SIB ALA CAN GRL

Comments:


15.1.4  Sparganium gramineum Georgi (1775), Bemerk. Reise 1: 232.

S S. friesii Laest. ex Beurl. (1854), Bot. Not. 1854: 136.

2n= 30 (2x).

2nD Lohammar in Löve & Löve (1942b Sweden).

G RFE

Comments:



(1) Is the name S. friesii used in any of our standard sources? It could be accepted because it has been in quite recent use (outside our sources) in N Europe. (Elven)
15.1.5  Sparganium natans L. (1753), Sp. Pl. 971.

S S. minimum Wallr. (1840), Erst. Beitr. Fl. Hercyn. 2: 297.

2n= 30 (2x).

2nD Löve & Löve (1975) list several counts, one Icelandic; Hämet-Ahti et al. (1984 Finl).

G ICE NOR? RUS ALA CAN

Comments:

(1) Kent gives p. 294 for the publication of Wallroth's S. minimum. Check. (Elven)
15.1.6  Sparganium hyperboreum Laest. ex Beurl. (1853), Öfvers. Kongl. Vet.-Akad. Förh. 9: 192.

S

2n= 30 (2x).



2nD Löve & Löve (1975) list several counts, most as arctic.

G ICE NOR RUS SIB RFE ALA CAN GRL

Comments:

16  Butomaceae (Draft: N.N. Tzvelev)

16.1  Butomus L. (1753), Sp. Pl. 372.
16.1.1  Butomus umbellatus L. (1753), Sp. Pl. 372.

S

2n= 16, 20, 22, 24, 26, 28, 30, 39, 40, 42.



2nD Löve & Löve (1975) list numerous 2n=26 counts and several 2n=39 counts, none as arctic; Hämet-Ahti et al. (1984 Finl 2n=26). See also Bolkhovskikh et al. (1969).

G RUS


Comments:

(1) Border occurrences in European Russia. (Elven)



17  Alismataceae (Draft: N.N. Tzvelev)

17.1  Alisma L. (1753), Sp. Pl. 342.
The Alisma plantago-aquatica aggregate (A. plantago-aquatica, A. triviale)

Comments:

(1) The Eurasian plant that reaches the Arctic is A. plantago-aquatica (subsp. plantago-aquatica). A North American parallel is variously treated as a species (A. triviale), subspecies (subsp. brevipes (Greene) Sam.) og variety (var. americanum Schult. & Schult. f.). It reaches the Arctic both on the SW side of Hudson Bay (York Factory) and im Seward Peninsula. It is recognised as species by Haynes & Hellquist (2000, Fl. N. Amer. 22) and that treatment is followed here. (Elven)


17.1.1  Alisma plantago-aquatica L. (1753), Sp. Pl. 342.

S A. plantago-aquatica L. sso. plantago-aquatica.

2n= 10, 12, 14 (2x), 16, 28.

2nD Löve & Löve (1975) list numerous non-arctic 2n=14 counts; Hämet-Ahti et al. (1984 Finl 2n=14). See also Bolkhovskikh et al. (1969).

G RUS SIB

Comments:


17.1.2  Alisma triviale Pursh (1814), Fl. Amer. Sept. 1: 252.

S A. plantago-aquatica L. subsp. brevipes (Greene) Sam. ***; A. plantago-aquatica L. var. americanum Schult. & Schult. fil. ***

2n= 28 (4x).

2nD Haynes & Hellquist (2000, Fl. N. Amer. 22, secondary counts)

G ALA CAN

Comments


(1) Given (for the collective species) for York Factory by Scoggan (1978) and for Seward Peninsula by Haynes & Hellquist (2000). (Elven)

17.2  Sagittaria L. (1753), Sp. Pl. 993.
17.2.1  Sagittaria cuneata Sheldon (1893), Bull. Torrey Bot. Club 20: 283.

S

2n= 22 (2x).



2nD Packer (1964 N Am); Löve & Ritchie (1966 Can).

G CAN


Comments:

(1) Border occurrences in the Mackenzie River delta and along the south shore of Hudson Bay (at least Fort Severn). (Elven)


17.2.2  Sagittaria natans Pallas (1776), Reise 3: 757.

S

2n= 22 (2x).



2nD Lohammar (1931a Sweden); Vaarama (Finl) etc.; Hämet-Ahti et al. (1984 Finl).

G SIB


Comments:

(1) Border occurrence in the Yamal-Gydan region. (Elven)



18  Hydrocharitaceae (Draft: R. Elven)

Comments:

(1) Family added to Tzvelev's draft. (Elven)

18.1  Elodea L.C. Richard in Michx. (1803), Fl. Bor. Amer. 1: 20.
18.1.1  Elodea canadensis L.C. Richard in Michx. (1803), Fl. Bor. Amer. 1: 20.

S

T [Described from Canada: near Montreal.]



2n= (1) 24 (2x). (2) 48 (4x), but see comment.

2nD (1) Heppell (1945 W Eur). (2) Skalinska & Pogan (1973 C Eur).

G CAN?

Comments:



(1) Acc. to Hultén & Fries (1986, map 94), this species approaches or reaches the Arctic on the south shore of Hudson Bay but Scoggan (1978) only reports it north to Reindeer Lake in Manitoba, 5754'N. Haynes (2000, Fl. N. Amer. 22) does not mark any arctic American sites in his map. The species, genus and family have therefore only very tentatively been added to Tzvelev's draft. Canadians must decide whether there are occurrences inside the Arctic as defined. (Elven)

(2) The 2n=48 number is more typical of E. nuttallii (Planchon) H. St. John, also spread in Europe. (Elven)



WARNING! Will be excluded if arctic occurrence in Canada is not confirmed.

19  Scheuchzeriaceae (Draft: N.N. Tzvelev)

19.1  Scheuchzeria L. (1753), Sp. Pl. 338.
19.1.1  Scheuchzeria palustris L. (1753), Sp. Pl. 338.

S

2n= 22 (2x).



2nD Löve & Löve (1975) list several non-arctic counts for the collective species.

Comments:

(1) The Eurasian race   subsp. palustris - does not seem to reach or even closely approach the Arctic. (Elven)
19.1.1.1  Scheuchzeria palustris L. subsp. americanus (Fern.) Hultén (1941), Lunds Univ. Årsskr., n.f., avd. 2, 37: 107.

B Scheuchzeria palustris L. var. americanus Fern. (1923), Rhodora 25: 178.

S

2n= 22 (2x).



2nD Löve & Ritchie (1966 Can).

G CAN?


Comments:

(1) The American entity is a major race and differs, according to Hultén (1962), in at least two presumed independent characters: longer fruits with a curved beak (1-2 mm long vs. more or less beakless in subsp. palustris) and larger seeds (4-5 mm vs. 3-4 mm). Status as subspecies is the obvious choice if these differences hold true. They are, however, doubted by Nienaber (2000, Fl. N. Amer. 22): "Variability in those characters, in specimens from both hemispheres, vitiates their worth for varietal distinction". (Elven)

(2) Another question is whether it reaches the Arctic as defined. Canadians must check. (Elven)

WARNING! Will be excluded if arctic occurrence in Canada not confirmed.

20  Juncaginaceae (Draft: N.N. Tzvelev)

20.1  Triglochin L. (1753), Sp. Pl. 338.

Comments:

(1) The arguments of Haynes & Hellquist (2000, Fl. N. Amer. 22) for treatment of Triglochin as feminine rather than neuter is accepted here as they are based on the current Code. (Elven)
20.1.1  Triglochin maritima L. (1753), Sp. Pl. 339.

S

2n= 12, 24, 30, 36, 48, 120.



2nD Löve & Löve (1975) list numerous 2n=48 counts; Hämet-Ahti et al. (1984 Finl 2n=48). See also Bolkhovskikh et al. (1969).

G ICE NOR RUS SIB ALA CAN

Comments:

(1) Hultén & Fries (1986 map 97) also map an occurrence in Russian Far East (East Chukotka). What about it? (Elven)


20.1.2  Triglochin palustris L. (1753), Sp. Pl. 338.

S

2n= (1) 24 (4x). (2) 24-26 (4x).



2nD (1) Löve & Löve (1975) list several counts, some as arctic; Hämet-Ahti et al. (1984 Finl); see also Bolkhovskikh et al. (1969). (2) Sokolovskaya (1963).

G ICE NOR RUS RFE ALA CAN GRL



Comments:


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